Queen of the San Diego Zoo Passes Peacefully at 141

The San Diego Zoo is mourning the loss of one of its most beloved residents. Gramma, a Galápagos tortoise who spent nearly a century charming visitors with her calm, quiet presence, has died at an estimated age of 141.

She was the zoo’s oldest living resident and a gentle reminder of nature’s quiet resilience.

Gramma passed away peacefully, according to zoo officials. While her exact birth year is unknown, records suggest she hatched sometime in the 1880s—decades before automobiles were common or radios filled the airwaves. She arrived at the San Diego Zoo around 1928, transferred from the Bronx Zoo as part of an early animal exchange program…

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